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Your test of manually controlling the lockup is a valid one. But if it's only happening in 4th lockup, under 45 mph, that's low RPM/moderate load, then I think the issue is with your tune. Try knocking a few degrees back on the timing, even if temporarily just for testing. You might need to do some limiting on your vacuum advance.

-Dave
 

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I'm saying it might -- I'm basing that on my experience with a mild 350 and a 700r4 lockup. I had a Holley Sniper on there controlling ignition and the default ignition map had a setting of 40 in certain areas which was too much, and I had the same kind of symptoms you describe. I lowered the timing in some of those areas and it helped. Granted, it was a lot easier to do that on EFI. But that is the experience I'm basing my comment/suggestion on.

-Dave
 
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